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That ability is "when the last defense counter is removed from this permanent, exile it, then you may cast it transformed without paying its mana cost." This means that if for any reason a battle enters the battlefield without defense counters, it will be put into the graveyard and its controller will not get the chance to exile it

Boros is enjoying something of a renaissance in Commander, with plenty of fun and interesting deck building opportunities

Two 2017 World Championships Deck (Golisodor - Naoto Suzuki)

Grading companies prefer them, and theyre safer for shipping

Draft B (on the right) is an evolution of the idea I originally had of drawing Arcanine on a mountain top